Mumble (Elijah Wood), the tap-dancing penguin, and Gloria (Alecia Moore {P!nk}), the love of his life, have a tiny son, Erik (EG Daily), who's reluctant to dance. He runs away and meets The Mighty Sven—a penguin who can fly! Sadly, Mumble realizes he can't compete with his son's new role model.

But new dangers are threatening the penguin nation, and Erik finds out just what his father's made of when Mumble brings not only the penguins together and gets them working—and dancing—but also creatures such as the tiny Krill and the giant Elephant Seals in an effort to save their home and way of life.